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William Eberhard


Méndez, Vivian and Eberhard, William G. 2014. Removal of genital plugs and insemination by males with normal and experimentally modified palps in Leucauge mariana (Araneae: Tetragnathidae). Journal of Arachnology, 42(2):284-292

Aisenberg, Anita, Barrantes, Gilbert and Eberhard, William G. 2014. Hairy kisses: tactile cheliceral courtship affects female mating decisions in Leucauge mariana (Araneae, Tetragnathidae). Doi: 10.1007/s00265-014-1844-2


Ang, Yuchen; Puniamoorthy, Jayanthi; Pont, Adrian C.; Bartak, Miroslav; Blanckenhorn, Wolf U.; Eberhard, William G.; Puniamoorthy, Nalini; Silva, Vera C.; Munari, Lorenzo; Meier, Rudolf. 2013. A plea for digital reference collections and other science-based digitization initiatives in taxonomy: Sepsidnet as exemplar. Systematic Entomology, 38(3): 637-644.


Eberhard, William G.; Hesselberg, Thomas. 2012. Cues that Spiders (Araneae: Araneidae, Tetragnathidae) Use to Build Orbs: Lapses in Attention to One Set of Cues because of Dissonance with Others?. Ethology, 118(7): 610-620.

Eberhard, William G.; Wcislo, William T.. 2012. Plenty of room at the bottom? Tiny animals solve problems of housing and maintaining oversized brains, shedding new light on nervous-system evolution. American Scientist, 100(3): 226-233..

Briceno, R. D.; Eberhard, William G.. 2012. Spiders avoid sticking to their webs: clever leg movements, branched drip-tip setae, and anti-adhesive surfaces. Naturwissenschaften, 99(4): 337-341..

Barrantes, Gilbert; Eberhard, William G.. 2012. Extreme Behavioral Adjustments by an Orb-Web Spider to Restricted Spaces. Ethology, 118(5): 438-449..


Eberhard, W. G. 2011. Experiments with genitalia: a commentary. Trends in Ecology and Evolution 26(1), 17-21. doi:10.1016/j.tree.2010.10.009

Briceño, R. D. & Eberhard, W. G. 2011. The hub as a launching platform: rapid movements of the spider Leucauge mariana (Araneae: Tetragnathidae) as it turns to attack prey. Journal of Arachnology, 39(1), 102–112.

Eberhard, W. G. & Wcislo, W. T. 2011. Morphological and behavioral consequences of physiological limitations in miniature arthropods. Advances in Insect Physiology.

Quesada, Rosannette; Triana, Emilia; Vargas, Gloria; Douglass, John K.; Seid, Marc A.; Niven, Jeremy E.; Eberhard, William G.; Wcislo, William T.. 2011. The allometry of CNS size and consequences of miniaturization in orb-weaving and cleptoparasitic spiders. Arthropod Structure & Development, 40(6): 521-529..

Eberhard, W. G. 2011. Are smaller animals behaviourally limited? Lack of clear constraints in miniature spiders. Animal Behaviour, 81(4), 813-823.


Eberhard, W. G. 2010. Possible functional significance of spigot placement on the spinnerets of spiders. Journal of Arachnology, 38(3), 407–414.

Eberhard, W.G. (2010) Cryptic Female Choice. In: Breed M.D. and Moore J., (eds.) Encyclopedia of Animal Behavior, volume 1, pp. 430-434 Oxford: Academic Press.

Barrantes, G. & Eberhard, W. G. 2010. Ontogeny repeats phylogeny in Steatoda and Latrodectus spiders. Journal of Arachnology, 38(3), 485–494.

Gonzaga, M., Sobczak, J., Penteado-Dias, A. & Eberhard, W. 2010. Modification of Nephila clavipes (Araneae Nephilidae) webs induced by the parasitoids Hymenoepimecis bicolor and H. robertsae (Hymenoptera Ichneumonidae). Ethology Ecology & Evolution, 22(2), 151–165.

Eberhard, W. G. 2010. Recovery of spiders from the effects of parasitic wasps: implications for fine-tuned mechanisms of manipulation. Animal Behaviour, 79(2), 375–383.

Eberhard, W. G. & Huber, B. A. 2010. Spider Genitalia. Pp. 249-284 in J. Leonard & A. Cordoba (eds.) The Evolution of Primary Sexual Characters in Animals.

Eberhard, W. G. 2010. Rapid divergent evolution of genitalia. Pp. 40-78 in J. Leonard & A. Cordoba (eds.) The Evolution of Primary Sexual Characters in Animals.

Moya, J., R. Quesada, G. Barrantes, WG Eberhard, I. Escalante, C. Esquivel, A. Rojas, E. Triana and A. Arias. 2010. Egg sac construction by folding dead leaves in Pozonia nigroventris and Micrathena sp. (Araneae: Araneidae). Journal of Arachnology, 38(2):371-373.

Briceño, R. D., Wegrzynek, D., Chinea-Cano, E., Eberhard, W. G. & dos Santos Rolo, T. 2010. Movements and morphology under sexual selection: tsetse fly genitalia. Ethology Ecology & Evolution, 22(4), 385–391.

Eberhard, W. G. 2010. New types of behavioral manipulation of host spiders by a parasitoid wasp. Psyche, Article ID 950614, 1-4.


Eberhard, W. G. 2009. Postcopulatory sexual selection: Darwin’s omission and its consequences. Proceedings of the National Academy of Sciences, 106(Suppl 1), 10025-10032.

Briceño, R. D. & Eberhard, W. G. 2009. Experimental modifications imply a stimulatory function for male tsetse fly genitalia, supporting cryptic female choice theory. Journal of Evolutionary Biology, 22(7), 1516–1525.

Peretti, A. V. & Eberhard, W. G. 2009. Cryptic female choice via sperm dumping favours male copulatory courtship in a spider. Journal of Evolutionary Biology, 23(2), 271–281.

Eberhard, W. G. 2009. Book Review: Art for art’s sake. Ethology, 115, 1-2.

William G. Eberhard. 2009. Evolution of genitalia – theories, evidence, and new directions. Genetica. 2009.

Eberhard, W. G., Rodriguez, R. L. & Polihronakis, M. 2009. Pitfalls in understanding the functional significance of genital allometry. Journal of Evolutionary Biology, 22(3), 435–445.

Briceño, R. D. & Eberhard, W. G. 2009. Experimental demonstration of possible cryptic female choice on male tsetse fly genitalia. Journal of Insect Physiology, 55(11), 989–996.

Aisenberg, A. & Eberhard, W. G. 2009. Female cooperation in plug formation in a spider: effects of male copulatory courtship. Behavioral Ecology, 20(6), 1236.

Eberhard, W. G. & Gelhaus, J. K. 2009. Genitalic stridulation during copulation in a species of crane fly, Tipula (Bellardina) sp. (Diptera: Tipulidae). Revista de Biología Tropical, 57(Suppl. 1), 251–256.

Eberhard, W. G. 2009. Static allometry and animal genitalia. Evolution, 63(1), 48–66.


Eberhard, W. G. 2008. Araneus expletus (Araneae, Araneidae): another stabilimentum that does not function to attract prey. Journal of Arachnology, 36(1), 191–194.

Barrantes, G., Eberhard, W. G. & Weng, J. L. 2008. Seasonal patterns of parasitism of the tropical spiders Theridion evexum (Araneae, Theridiidae) and Allocyclosa bifurca (Araneae, Araneidae) by the wasps Zatypota petronae and Polysphincta gutfreundi (Hymenoptera, Ichneumonidae). Revista de Biología Tropical, 56(2), 749–754.

Eberhard, W. G., Agnarsson, I. & Levi, H. W. 2008. Web forms and the phylogeny of theridiid spiders (Araneae: Theridiidae): chaos from order. Systematics and Biodiversity, 6(4), 415–475.

Eberhard, W. G., Barrantes, G. & Madrigal-Brenes, R. 2008. Vestiges of an orb-weaving ancestor? The “biogenetic law” and ontogenetic changes in the webs and building behavior of the black widow spider Latrodectus geometricus (Araneae Theridiidae). Ethology Ecology & Evolution, 20(3), 211–244.


Jörger, K. M. & Eberhard, W. G. 2006. Web construction and modification by Achaearanea tesselata (Araneae, Theridiidae). Journal of Arachnology, 34(3), 511–523.

Briceño, R. D., Eberhard, W. G. & Robinson, A. S. 2007. Copulation behaviour of Glossina pallidipes (Diptera: Muscidae) outside and inside the female, with a discussion of genitalic evolution. Bulletin of Entomological Research, 97(5), 471–488.

Eberhard, W. G. 2007. Miniaturized orb-weaving spiders: behavioural precision is not limited by small size. Proceedings of the Royal Society B: Biological Sciences, 274(1622), 2203-2209.

Baena, ML; Eberhard, W. G. 2007. Appearances deceive: female "resistance" behaviour in a sepsid fly is not a test of male ability to hold on. Ethology Ecology & Evolution. 19 (1) pp. 27-50.

Briceño, R. D., Eberhard, W. G. & Shelly, T. 2007. Male courtship behavior in Ceratitis capitata (Diptera: Tephritidae) that have received aromatherapy with ginger root oil. Florida Entomologist, 90(1), 175–179.

Eberhard, W. G. 2007. Stabilimenta of Philoponella vicina (Araneae: Uloboridae) and Gasteracantha cancriformis (Araneae: Araneidae): evidence against a prey attractant function. Biotropica, 39(2), 216–220.

Eberhard, W. G. 2007. Swarming behavior of two parasitic wasps, Leurus caeruliventris (Hymenoptera: Ichneumonidae) and Conura sp.(Hymenoptera: Chalcididae), at the same site. Journal of the Kansas Entomological Society, 80(3), 255–258.

Barrantes, G. & Eberhard, W. G. 2007. The evolution of prey-wrapping behaviour in spiders. Journal of Natural History, 41(25-28), 1631–1658.

Briceño, R. D., Eberhard, W. G., Vilardi, J., Cayol, J. P. & Shelly, T. 2007. Courtship behavior of different wild strains of Ceratitis capitata (Diptera: Tephritidae). Florida Entomologist, 90(1), 15–18.


Eberhard, W. G. 2006. Sexually antagonistic coevolution in insects is associated with only limited morphological diversity. Journal of Evolutionary Biology. 19(3):657-681.

Eberhard, W. G., Barrantes, G. & Weng, J. L. 2006. Tie them up tight: wrapping by Philoponella vicina spiders breaks, compresses and sometimes kills their prey. Naturwissenschaften 93(5), 251-254.

Daniel Briceño, William Eberhard, Juan Vilardi, Jean-Pierre Cayol and Todd Shelly. 2006. Courtship Behavior of Different Wild Strains of Ceratitis Capitata (Diptera: Tephritidae).

Flowers, W. & Eberhard, W. G. 2006. Fitting together: copulatory linking in some Neotropical Chrysomeloidea. Revista de Biología Tropical, 54(3), 829–842

Eberhard, W. G. 2006. Sexually Reversed Copulatory courtship roles and possible nuptial feeding in the soldier beetle Ditemnus acantholabus (Champion 1915) (Coleoptera: Cantharidae). Journal of the Kansas Entomological Society, 79(1), 13–22.

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