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William Eberhard


Eberhard, William G. 2018. The Webs of Neoantistea riparia (Araneae: Hahniidae): Are Dew Drops Helpful in Prey Capture?. Arachnology. 17 (9):485–490.

Eberhard, William G. 2018. Modular patterns in behavioural evolution: webs derived from orbs. Behaviour. 155 (6):531–566.

Eberhard, William G., Rodríguez, Rafael Lucas, Huber, Bernhard A., Speck, Bretta, Miller, Henry, Buzatto, Bruno A., and Machado, Glauco. 2018. Sexual Selection and Static Allometry: The Importance of Function. The Quarterly review of biology. 93 (3):207–250.


Eberhard, William G. 2017. Cryptic Female Choice Revisited: A Response to Firman et al.. Trends in Ecology & Evolution. 32 (11):805–806.

Eberhard, William G. and Hazzi, Nicolas A. 2017. Web building and prey wrapping behavior of Aglaoctenus castaneus (Araneae: Lycosidae: Sosippinae). Journal of Arachnology. 45 (2):177–197.

Eberhard, William G. 2017. How orb-weavers find and grasp silk lines. Journal of Arachnology. 45 (2):145–151.


Briceno, R. D., Eberhard, William G., Chinea-Cano, E., Wegrzynek, D., and dos Santos Rolo, T. 2016. Species-specific differences in the behavior of male tsetse fly genitalia hidden in the female during copulation. Ethology Ecology & Evolution. 28 (1):53–76.


Eberhard, William G. and Barrantes, Gilbert. 2015. Cues guiding uloborid construction behavior support orb web monophyly. Journal of Arachnology. 43 (3):371–387.

Eberhard, William G. 2015. How Micrathena duodecimspinosa (Araneae: Araneidae) uses the elasticity of her dragline to hide her egg sac. Journal of Arachnology. 43 (3):417–418.

Aisenberg, Anita, Barrantes, Gilbert, and Eberhard, William G. 2015. Post-copulatory Sexual Selection in Two Tropical Orb-weaving Leucauge Spiders. in Cryptic female choice in arthropods: Patterns, mechanisms and prospects, edited by Peretti, Alfredo V. and Aisenberg, Anita., 79–108. Germany: Springer.


Eberhard, William G., Pacheco, Jessica, Carrasco-Rueda, Farah, Christopher, Yuliana, Gonzalez, Cely, Urbina, Hector, and Blackwell, Meredith. 2014. Zombie bugs? The fungus Purpureocillium sp. may manipulate the behavior of its host bug Edessa rufomarginata. Mycologia. 106 (6):1065–1072.

Aisenberg, Anita, Barrantes, Gilbert and Eberhard, William G. 2015. Hairy kisses: tactile cheliceral courtship affects female mating decisions in Leucauge mariana (Araneae, Tetragnathidae). Doi: 10.1007/s00265-014-1844-2

Eberhard, William G. 2014. A new view of orb webs: multiple trap designs in a single structure. Biological Journal of the Linnean Society. 111 (2):437–449.

Méndez, Vivian and Eberhard, William G. 2014. Removal of genital plugs and insemination by males with normal and experimentally modified palps in Leucauge mariana (Araneae: Tetragnathidae). Journal of Arachnology, 42(2):284-292


Ang, Yuchen; Puniamoorthy, Jayanthi; Pont, Adrian C.; Bartak, Miroslav; Blanckenhorn, Wolf U.; Eberhard, William G.; Puniamoorthy, Nalini; Silva, Vera C.; Munari, Lorenzo; Meier, Rudolf. 2013. A plea for digital reference collections and other science-based digitization initiatives in taxonomy: Sepsidnet as exemplar. Systematic Entomology, 38(3): 637-644.


Eberhard, William G.; Wcislo, William T.. 2012. Plenty of room at the bottom? Tiny animals solve problems of housing and maintaining oversized brains, shedding new light on nervous-system evolution. American Scientist, 100(3): 226-233..

Briceno, R. D.; Eberhard, William G.. 2012. Spiders avoid sticking to their webs: clever leg movements, branched drip-tip setae, and anti-adhesive surfaces. Naturwissenschaften, 99(4): 337-341..

Barrantes, Gilbert; Eberhard, William G.. 2012. Extreme Behavioral Adjustments by an Orb-Web Spider to Restricted Spaces. Ethology, 118(5): 438-449..

Eberhard, William G.; Hesselberg, Thomas. 2012. Cues that Spiders (Araneae: Araneidae, Tetragnathidae) Use to Build Orbs: Lapses in Attention to One Set of Cues because of Dissonance with Others?. Ethology, 118(7): 610-620.


Briceño, R. D. & Eberhard, W. G. 2011. The hub as a launching platform: rapid movements of the spider Leucauge mariana (Araneae: Tetragnathidae) as it turns to attack prey. Journal of Arachnology, 39(1), 102–112.

Eberhard, W. G. & Wcislo, W. T. 2011. Morphological and behavioral consequences of physiological limitations in miniature arthropods. Advances in Insect Physiology.

Quesada, Rosannette; Triana, Emilia; Vargas, Gloria; Douglass, John K.; Seid, Marc A.; Niven, Jeremy E.; Eberhard, William G.; Wcislo, William T.. 2011. The allometry of CNS size and consequences of miniaturization in orb-weaving and cleptoparasitic spiders. Arthropod Structure & Development, 40(6): 521-529..

Eberhard, W. G. 2011. Are smaller animals behaviourally limited? Lack of clear constraints in miniature spiders. Animal Behaviour, 81(4), 813-823.

Eberhard, W. G. 2011. Experiments with genitalia: a commentary. Trends in Ecology and Evolution 26(1), 17-21. doi:10.1016/j.tree.2010.10.009


Gonzaga, M., Sobczak, J., Penteado-Dias, A. & Eberhard, W. 2010. Modification of Nephila clavipes (Araneae Nephilidae) webs induced by the parasitoids Hymenoepimecis bicolor and H. robertsae (Hymenoptera Ichneumonidae). Ethology Ecology & Evolution, 22(2), 151–165.

Eberhard, W. G. 2010. Recovery of spiders from the effects of parasitic wasps: implications for fine-tuned mechanisms of manipulation. Animal Behaviour, 79(2), 375–383.

Eberhard, W. G. & Huber, B. A. 2010. Spider Genitalia. Pp. 249-284 in J. Leonard & A. Cordoba (eds.) The Evolution of Primary Sexual Characters in Animals.

Eberhard, W. G. 2010. Rapid divergent evolution of genitalia. Pp. 40-78 in J. Leonard & A. Cordoba (eds.) The Evolution of Primary Sexual Characters in Animals.

Moya, J., R. Quesada, G. Barrantes, WG Eberhard, I. Escalante, C. Esquivel, A. Rojas, E. Triana and A. Arias. 2010. Egg sac construction by folding dead leaves in Pozonia nigroventris and Micrathena sp. (Araneae: Araneidae). Journal of Arachnology, 38(2):371-373.

Briceño, R. D., Wegrzynek, D., Chinea-Cano, E., Eberhard, W. G. & dos Santos Rolo, T. 2010. Movements and morphology under sexual selection: tsetse fly genitalia. Ethology Ecology & Evolution, 22(4), 385–391.

Eberhard, W. G. 2010. New types of behavioral manipulation of host spiders by a parasitoid wasp. Psyche, Article ID 950614, 1-4.

Eberhard, W. G. 2010. Possible functional significance of spigot placement on the spinnerets of spiders. Journal of Arachnology, 38(3), 407–414.

Eberhard, W.G. (2010) Cryptic Female Choice. In: Breed M.D. and Moore J., (eds.) Encyclopedia of Animal Behavior, volume 1, pp. 430-434 Oxford: Academic Press.

Barrantes, G. & Eberhard, W. G. 2010. Ontogeny repeats phylogeny in Steatoda and Latrodectus spiders. Journal of Arachnology, 38(3), 485–494.


Peretti, A. V. & Eberhard, W. G. 2009. Cryptic female choice via sperm dumping favours male copulatory courtship in a spider. Journal of Evolutionary Biology, 23(2), 271–281.

Eberhard, W. G. 2009. Book Review: Art for art’s sake. Ethology, 115, 1-2.

William G. Eberhard. 2009. Evolution of genitalia – theories, evidence, and new directions. Genetica. 2009.

Eberhard, W. G., Rodriguez, R. L. & Polihronakis, M. 2009. Pitfalls in understanding the functional significance of genital allometry. Journal of Evolutionary Biology, 22(3), 435–445.

Briceño, R. D. & Eberhard, W. G. 2009. Experimental demonstration of possible cryptic female choice on male tsetse fly genitalia. Journal of Insect Physiology, 55(11), 989–996.

Aisenberg, A. & Eberhard, W. G. 2009. Female cooperation in plug formation in a spider: effects of male copulatory courtship. Behavioral Ecology, 20(6), 1236.

Eberhard, W. G. & Gelhaus, J. K. 2009. Genitalic stridulation during copulation in a species of crane fly, Tipula (Bellardina) sp. (Diptera: Tipulidae). Revista de Biología Tropical, 57(Suppl. 1), 251–256.

Eberhard, W. G. 2009. Static allometry and animal genitalia. Evolution, 63(1), 48–66.

Eberhard, W. G. 2009. Postcopulatory sexual selection: Darwin’s omission and its consequences. Proceedings of the National Academy of Sciences, 106(Suppl 1), 10025-10032.

Briceño, R. D. & Eberhard, W. G. 2009. Experimental modifications imply a stimulatory function for male tsetse fly genitalia, supporting cryptic female choice theory. Journal of Evolutionary Biology, 22(7), 1516–1525.


Eberhard, W. G., Agnarsson, I. & Levi, H. W. 2008. Web forms and the phylogeny of theridiid spiders (Araneae: Theridiidae): chaos from order. Systematics and Biodiversity, 6(4), 415–475.

Eberhard, W. G., Barrantes, G. & Madrigal-Brenes, R. 2008. Vestiges of an orb-weaving ancestor? The “biogenetic law” and ontogenetic changes in the webs and building behavior of the black widow spider Latrodectus geometricus (Araneae Theridiidae). Ethology Ecology & Evolution, 20(3), 211–244.

Eberhard, W. G. 2008. Araneus expletus (Araneae, Araneidae): another stabilimentum that does not function to attract prey. Journal of Arachnology, 36(1), 191–194.

Barrantes, G., Eberhard, W. G. & Weng, J. L. 2008. Seasonal patterns of parasitism of the tropical spiders Theridion evexum (Araneae, Theridiidae) and Allocyclosa bifurca (Araneae, Araneidae) by the wasps Zatypota petronae and Polysphincta gutfreundi (Hymenoptera, Ichneumonidae). Revista de Biología Tropical, 56(2), 749–754.


Eberhard, W. G. 2007. Miniaturized orb-weaving spiders: behavioural precision is not limited by small size. Proceedings of the Royal Society B: Biological Sciences, 274(1622), 2203-2209.

Baena, ML; Eberhard, W. G. 2007. Appearances deceive: female "resistance" behaviour in a sepsid fly is not a test of male ability to hold on. Ethology Ecology & Evolution. 19 (1) pp. 27-50.

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