STRI Coral Reef

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Rachel Page


Carter G, Wilkinson G, Page RA. 2017. Food-sharing vampire bats are more nepotistic under conditions of perceived risk. Behavioral Ecology. doi:10.1093/beheco/arx006.

Hemingway, C., Ryan, M.J., Page, R.A. 2017. Rationality in decision-making in the fringe-lipped bat, Trachops cirrhosus. Behavioral Ecology and Sociobiology. 71: 94. doi:10.1007/s00265-017-2321-5.

ter Hofstede, H.M., Voigt-Heucke, S.L., Lang, A., Römer, H., Page, R.A., Faure, P.A., Dechmann, D.K.N. 2017. Revisiting adaptations of Neotropical katydids (Orthoptera: Tettigoniidae) to gleaning bat predation. Neotropical Biodiversity. 3: 41-49. doi: 10.1080/23766808.2016.1272314.


Jones, P.L., Page, R.A., Ratcliffe, J.M. In press. To scream or to listen? Prey detection and discrimination in animal-eating bats. In: Bat Bioacoustics (volume editors: B. Fenton and A. Grinnell; series editor: A. Popper). Springer.

Stange, N., Page, R.A., Ryan, M.J., Taylor, R.C. In press. Interactions between complex multisensory signal components result in unexpected mate choice responses. Animal Behaviour.

Vencl, F.V., Ottens, K., Dixon, M.M., Candler, S., Bernal, X.E., Estrada, C., Page, R.A. 2016. Pyrazine emission by a tropical firefly: an example of chemical aposematism? Biotropica.

Page, R.A., Jones, P.L. In press. Overcoming sensory uncertainty: factors affecting foraging decisions in frog-eating bats. In: Perception and Cognition in Animal Communication (volume editors, M.A. Bee and C.T. Miller), in the book series Animal Signals and Communication (series editors: P.K. McGregor and V.M. Janik). Springer.

Vencl, F.V., Ottens, K., Dixon, M.M., Candler, S., Bernal, X.E., Estrada, C., Page, R.A. 2016. Pyrazine emission by a tropical firefly: an example of chemical aposematism? Biotropica. 48: 645–655. doi:10.1111/btp.12336.

Ramakers J.J.C, Dechmann D.K.N., Page R.A., O’Mara M.T. (2016) Frugivorous bats prefer information from novel social partners. Animal Behaviour. 116: 83-87.

Symes, L.B., Page, R.A., ter Hofstede, H.M. In press. Interspecific variation in call timing by Neotropical forest katydids. Behavioral Ecology and Sociobiology.

Gomes, D. G. E., Page, R. A., Geipel, I., Taylor, R. C., Ryan, M. J., Halfwerk, W. 2016. Bats perceptually weight prey cues across sensory systems when hunting in noise. Science. 353: 1277-1280. doi: 10.1126/science.aaf7934.

Halfwerk, W., Guerra, M.A., Lea, A.M., Page, R.A., Ryan, M.J. 2016. Amplitude regulation in frogs reveals the ancestral state of the Lombard effect. Behavioral Ecology. 27: 669-676. doi:10.1093/beheco/arv204.

Ripperger, S., Josic, D., Hierold, M., Koelpin, A., Weigel, R., Hartmann, M., Page, R.A., Mayer, F. 2016. Automated proximity sensing in small vertebrates: design of miniaturized sensor nodes and first field tests in bats. Ecology and Evolution. 6: 2179–2189.

Trillo, P.A., Bernal, X.E., Caldwell, M., Halfwerk, W., Owens, M., Page, R.A. 2016. Collateral damage or a shadow of safety? The effects of signaling neighbors on the risks of parasitism and predation. Proceedings of the Royal Society Series B. 283: 20160343.


Rhebergen, F., Taylor, R.C., Ryan, M.J., Page, R.A., Halfwerk, W. 2015. Multimodal cues improve prey localisation under complex environmental conditions. Proceedings of the Royal Society Series B. 282: 20151403.

Falk, J.J., ter Hofstede, H.M., Jones, P.L., Dixon, M.M., Faure, P.A., Kalko, E.K.V., Page, R.A. 2015. Sensory-based niche partitioning in a multiple predator-multiple prey community. Proceedings of the Royal Society Series B. 282: 20150520.

Bulbert, Matthew W., Page, Rachel A. and Bernal, Ximena E. 2015. Danger comes from all fronts: predator-dependent escape tactics of túngara frogs. PloS One, 10(4): 1-12. doi:10.1371/journal.pone.0120546

Fugère, V., O'Mara, M.T., Page, R.A. 2015. Perceptual bias does not explain preference for prey call adornment in the frog-eating bat. Behavioral Ecology and Sociobiology. DOI 10.1007/s00265-015-1949-2.

Bader, E., Jung, K., Kalko, E.K.V., Page, R.A., Rodriguez, R., Sattler, T. 2015. Mobility explains the response of aerial insectivorous bats to anthropogenic habitat change in the Neotropics. Biological Conservation. 186: 97–106.

Stockmaier, S., Dechmann, K.N., Page, R.A., O’Mara, M.T. 2015. No fever and leukocytosis in response to a lipopolysaccaride challenge in an insectivorous bat. Biology Letters. 11: 20150576.


Halfwerk, W., Jones, P.L., Taylor, R.C., Ryan, M.J., Page, R.A. 2014. Risky ripples allow bats and frogs to eavesdrop on a multisensory sexual display. Science. 342: 413-416, DOI 10.1126/science.1244812.

O’Mara, M.T., Dechmann, D.D, Page, R.A. 2014. Frugivorous bats evaluate the quality of social information when choosing novel foods. Behavioral Ecology. 1–7. doi:10.1093/beheco/aru120.

Clarin, T.M.A., Borissov, I., Page, R.A., Ratcliffe, J.M., Siemers, B.M. 2014. Social learning within and across species: information transfer in mouse-eared bats. Canadian Journal of Zoology.129-139. DOI 10.1139/cjz-2013-0211.

Halfwerk, Wouter, Page, Rachel A, Taylor, Ryan C, Wilson, Preston S and Ryan, Michael J. 2014. Crossmodal Comparisons of Signal Components Allow for Relative-Distance Assessment. Current Biology, 24(15): 1751-1755. doi:10.1016/j.cub.2014.05.068

Page, R.A., Ryan, M.J., Bernal, X.E. 2014. Be loved, be prey, be eaten. In: Animal Behavior, vol 3. Case Studies: Integration and Application of Animal Behavior (ed., K. Yasukawa), New York: Praeger. pp. 123-154.

Jones, P.L., Ryan, M.J., Page, R.A. 2014. Population and seasonal variation in response to prey calls by an eavesdropping bat. Behavioral Ecology and Sociobiology. DOI 10.1007/s00265-013-1675-6.

Halfwerk, W., Dixon, M.M., Ottens, K., Taylor, R.C., Ryan, M.J., Page, R.A., Jones, P.L. 2014. Risks of multimodal signaling: bat predators attend to dynamic motion in frog sexual displays. Journal of Experimental Biology. 217: 3038-3044. DOI 10.1242/jeb.107482.


Surlykke, Annemarie; Jakobsen, Lasse; Kalko, Elisabeth and Page, Rachel A. 2013. Echolocation intensity and directionality of perching and flying fringe-lipped bats, Trachops cirrhosis (Phyllostomidae). Frontiers in Physiology, vol. 4, art. 143 pp. 1-9

Jones, P.L., Farris, H.E., Ryan, M.J., Page, R.A. 2013. Do frog-eating bats perceptually bind the complex components of frog calls? Journal of Comparative Physiology A. 199: 279-283.

Jones, P.L., Ryan, M.J., Page, R.A. 2013. When to approach novel prey cues? Social learning strategies in frog-eating bats. Proceedings of the Royal Society B. 280: 20132330,

Clarin, Theresa M. A.; Ruczynski, Ireneusz; Page, Rachel A. and Siemers, Bjorn M. 2013. Foranging ecology predicts learning performance in insectivorous bats. PlosOne, vol. 8 (6) pp. 1-12


Page, R.A., von Merten, S. (equal contribution), Siemers, B.M. 2012. Associative memory or algorithmic search: a comparative study on learning strategies of bats and shrews. Animal Cognition.

Page, R.A., Schnelle, T., Kalko, E.K.V., Bunge, T., Bernal, X.E. 2012. Reassessment of prey through sequential use of multiple sensory cues by an eavesdropping bat. Naturwissenschaften 99: 505-509.


Jones, P.L., Page, R.A., Hartbauer, M., Siemers, B.M. 2011. Behavioral evidence for eavesdropping on prey song in two Palearctic sibling bat species. Behavioral Ecology and Sociobiology, 65: 333-340.

Akre, K.L., Farris, H.E., Lea, A.M., Page, R.A., Ryan, M.J. 2011. Signal perception in frogs and bats and the evolution of mating signals. Science 333: 751-752


Siemers, B.M., Page R.A. 2009. Behavioral studies of bats in captivity: Methodology, training, and experimental design. In: Thomas H. Kunz;Stuart Parsons (Ed.), Ecological and behavioral methods for the study of bats: . Baltimore, Maryland: The John Hopkins Press.

Bernal, X.E., Page, R.A., Ryan, M.J., Argo, T.F., and Wilson, P.S. 2009. Acoustic radiation patterns of mating calls of the túngara frog (Physalaemus pustulosus): implications for multiple receivers. Journal of the Acoustical Society of America. 126: 2757-2767.


Page, R.A. and M.J. Ryan. 2008. The effect of signal complexity on localization performance in bats that localize frog calls. Animal Behaviour 76: 761-769.


Page, R.A. 2007. Prey-predator communication: for your sensors only. Dispatch for Current Biology 17: R965-R966.

Bernal, X.E., Page, R.A., Rand, A.S., and Ryan, M.J. 2007. Cues for eavesdroppers: do frog calls indicate prey density and quality? American Naturalist 169: 409-415.


Page, R.A. and M.J. Ryan. 2006. Social transmission of novel foraging behavior in bats: frog calls and their referents. Current Biology, 16: 1201-1205.

Page, R.A. and X.E. Bernal. 2006. Túngara frogs. Quick guide for Current Biology 16: R979-980.


Page, R.A. and Ryan, M.J. 2005. Flexibility in assessment of prey cues: frog-eating bats and frog calls. Proceedings of the Royal Society Series B 272: 841-847.


Gabor, C.R. and Page, R.A. 2003. Female preference for large males in sailfin mollies, Poecilia latipinna: the importance of predation pressure and reproductive status. Acta Ethologica 6: 7-12.

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