Lessios, H.A. 1992. Testing electrophoretic data for agreement with Hardy-Weinberg expectations. Marine Biology, 112:517-523

Lessios, H.A. and J.S. Pearse. 1996. Hybridization and introgression between Indo-Pacific species of Diadema. Marine Biology, 126:715-723.

Lessios, H.A. . 1985b. Genetic consequences of mass mortality in the Caribbean sea urchin Diadema antillarum. Proceedings, 5th International Congress on Coral Reefs, 4:119-132

Lessios, H.A. 1981. Evidencia preliminar de una duplicacion del gen en la evolucion de la clase Echinoidea. Revista Medica, 6:46-55.

Lessios, H.A. and J. R. Weinberg 1993. Migration, gene flow and reproductive isolation between and within morphotypes of the isopod Excirolana in two oceans. Heredity 71:561-573

B. Stockley, A. B. Smith, T. Littlewood, H. A. Lessios, and J.A. Mackenzie-Dodds. 2005. Phylogenetic relationships of spatangoid sea urchins (Echinoidea): taxon sampling density and congruence between morphological and molecular estimates. Zoologica Scripta, 34:447-468

H.A. Lessios. 1987. Temporal and spatial variation in egg size of thirteen Panamanian echinoids. Journal of Experimental Marine Biology and Ecology, 114:217-239

H.A. Lessios. 1991.Presence and absence of monthly reproductive rhythms among eight Caribbean echinoids off the coast of Panama. Journal of Experimental Marine Biology and Ecology, 153:27-47

K.S. Zigler, M.A. McCartney, D. R. Levitan and H. A. Lessios. 2005. Sea urchin bindin divergence predicts gamete compatibility. Evolution 59:2399-2404

Lessios, H.A. 2005. Echinoids of the Pacific Waters of Panama:
Status of knowledge and new records. Rev. Biol. Tropical 53 (sup. 3):147-170