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heckadon-Moreno, Stanley 2017. Francisco Herrera y las represas de Fortuna y Changuinola. Epocas, 32(10): 10-11..


Bagley, Justin C., Alda, Fernando, Breitman, M. F., Bermingham, Eldredge, van, den Berghe and Johnson, Jerald B. 2015. Assessing Species Boundaries Using Multilocus Species Delimitation in a Morphologically Conserved Group of Neotropical Freshwater Fishes, the Poecilia sphenops Species Complex (Poeciliidae). PloS One, 10(4)

Heineman, Katherine D., Caballero, Pedro, Morris, Arturo, Velasquez, Carmen, Serrano, Kiria, Ramos, Nelly, Gonzalez, Jonathan, Mayorga, Luis, Corre, Marife D. and Dalling, James W. 2015. Variation in Canopy Litterfall Along a Precipitation and Soil Fertility Gradient in a Panamanian Lower Montane Forest. Biotropica, 47(3): 300-309..
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Rosindell, James, Hubbell, Stephen P., He, Fangliang, Harmon, Luke J. and Etienne, Rampal S. 2012. The case for ecological neutral theory. Trends in Ecology and Evolution, 27(4): 203-208..
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Jackson, Jeremy B. C. 2011. Response by Jeremy B.C. Jackson. Journal of Paleontology, 85(3): 601-602..

Ribeiro Mello, Marco Aurelio, Darcie Marquitti, Flavia Maria, Guimarães, Paulo R., Jr., Kalko, Elisabeth Klara Viktoria, Jordano, Pedro and Martinez de Aguiar, Marcus Aloizio 2011. The Missing Part of Seed Dispersal Networks: Structure and Robustness of Bat-Fruit Interactions. PloS One, 6(2): e17395..
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Schafer, Priska, Fortunato, Helena, Bader, Beate, Liebetrau, Volker, Bauch, Thorsten and Reijmer, John J. G. 2011. Growth rates and carbonate production by coralline red algae in upwelling and non-upwelling settings along the Pacific coast of Panama. Palaios, 26(7): 420-432..


Andersen, Kelly M., Turner, Benjamin L. and Dalling, James W. 2010. Soil-based habitat partitioning in understorey palms in lower montane tropical forests. Journal of Biogeography, 37(2): 278-292..
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Dalling, James W., Morris Valdes, Arturo, Caballero Ruiz, Pedro, Andersen, Kelly and Turner, Benjamin L. 2010. La dinámica del bosque de montaña en la Reserva Forestal de Fortuna, Chiriquí. Tecnociencia, 12(1): 119-135..
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Deichmann, Jessica L., Williamson, G. Bruce, Lima, Albertina P. and Allmon, Warren D. 2010. A note on amphibian decline in a central Amazonian lowland forest. Biodiversity and Conservation, 19(12): 3619-3627..


Fortunato, Helena M. and Schaefer, Priska 2009. Coralline algae as carbonate producers and habitat providers on the Eastern Pacific coast of Panama: preliminary assessment. Neues Jahrbuch für Geologie und Paläontologie. Abhandlungen, 253: 145-161..

Introíni, G. O., Magalhães, C. A., Fortunato, Helena M. and Recco-Pimentel, S. M. 2009. Comparison of the spermatozoan morphology of Isognomon bicolor and Isognomon alatus (Mollusca, Bivalvia, Isognomonidae). Tissue and Cell, 41(1): 67-74..


Fortunato, Helena M. 2008. The Central American land bridge: evolution at work. Schriften des Naturwissenschaftlichen Vereins für Schleswig-Holstein, 70: 56-72..

Robertson, D. Ross and Smith-Vaniz, William F. 2008. Rotenone: An essential but demonized tool for assessing marine fish diversity. Bioscience, 58(2): 165-170..
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Fortunato, H. 2007. Phylogenetic relationships of the columbellid taxa Cotonopsis and Cosmioconcha (Neogastropoda : Buccinoidea : Columbelfidae). American Malacological Bulletin, 23(1-2): 33-42..
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Fortunato, Helena M. 2007. Naticid gastropod predation in the Gatun Formation (late Middle Miocene), Panama: preliminary assessment. Paleontologische Zeitschrift, 81(3): 356-364..

O'Dea, Aaron, Jackson, Jeremy B. C., Fortunato, Helena M., Smith, J. Travis, D'Croz, Luis, Johnson, Kenneth G. and Todd, Jonathan A. 2007. Environmental change preceded Caribbean extinction by 2 million years. Proceedings of the National Academy of Sciences of the United States of America, 104(13): 5501-5506..
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Anker, Arthur, Ahyong, Shane T., Noel, Pierre Y. and Palmer, A. R. 2006. Morphological phylogeny of alpheid shrimps: Parallel preadaptation and the origin of a key morphological innovation, the snapping claw. Evolution, 60(12): 2507-2528..
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Fortunato, Helena M. 2004. Central American Neogene and Recent Marine Molluscan Diversity: An Overview for Conservation Purposes. Journal of Conchology, : 109-121..

Fortunato, Helena M. 2004. Reproductive Strategies in Gastropods Across the Panamá Seaway. Invertebrate Reproduction and Development, (2-3): 139-148..

Fortunato, Helena M. 2004. Gastropod Predation on the Panamanian Caribbean Coast. , : 479..

O'Dea, Aaron, Herrera-Cubilla, Amalia, Fortunato, Helena M. and Jackson, Jeremy B. C. 2004. Life history variation in cupuladriid bryozoans from either side of the Isthmus of Panamá. Marine Ecology Progress Series, 280: 145-161..
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Fortunato, Helena M. 2003. Reproduction and Larval Development of the Strombina-Group (Buccinoidea: Columbellidae) and Related Gastropods: Testing the Use of the Larval Shell for Inference of Development in Fossil Species. Bolettino Malacologico, 38(Suppl 4): 111-126..

Fortunato, Helena M. 2003. Evolutionary Paleontology and Informatics: the Neogene Marine Biota of Tropical America (NMITA) Database. , : 591-601..

Fortunato, Helena M. 2003. Habitat Change and Biodiversity Maintenance: Two Faces of the Same Process?. , : 38..
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Fortunato, Helena M. 2003. Foregut and Reproductive Tract Anatomy of the Strombina-Group (Neogastropoda: Columbellidae). Iberus, (2): 109-122..

Fortunato, Helena M. 2003. Biogeography and the Tempo of Speciation in Strombinid Gastropods. , : 259..
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Fortunato, Helena M. 2002. Foregut and Reproductive Tract Anatomy of the Strombina-Group (Neogastropoda: Columbellidae): A Preliminary Report. , : 135..

Fortunato, Helena M. 2002. The Systematic Position of Strombina (Cotonopsis) Lindae Petuch, 1988 (Gastropoda: Columbellidae). Nautilus, (2): 59-61..

Fortunato, Helena M., Gruber, Karl F. and Bermingham, Eldredge 2002. Evolution of the Strombina-Group (Neogastropoda: Columbellidae: Evidence From the Mitochondrial Data Compared With Morphology and the Fossil Record. , : 111..

Todd, Jonathan A., Jackson, Jeremy B. C., Johnson, Kenneth G., Fortunato, Helena M., Heitz, A., Alvarez F., Marcos A. and Jung, Peter 2002. The Ecology of Extinction: Molluscan Feeding and Faunal Turnover in the Caribbean Neogene. Proceedings of the Royal Society B: Biological Sciences, 269(1491): 571-578..
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Fortunato, Helena M. 2001. Los Moluscos y La Historia Natural De Panamá. , : 26 -31..

Fortunato, Helena M. 2001. Constraints in Development Modes: Adaptation of History. , : 104..

Fortunato, Helena M., Jackson, Jeremy B. C., Todd, Jonathan A., Alvarez F., Marcos A., Heitz, K. and Jung, Peter 2001. Molluscan Diversity in Tropical America Oceans: An Overview. , : 105..

Todd, Jonathan A., Jackson, Jeremy B. C., Johnson, Kenneth G., Fortunato, Helena M., Heitz, A. and Jung, Peter 2001. The Ecology of Extinction: Molluscan Food Webs Implicate Plummeting Nutrient Supply in Plio-Pleistocene Caribbean Faunal Turnover. , : 34 ..

Todd, Jonathan A., Jackson, Jeremy B. C., Johnson, Kenneth G., Fortunato, Helena M., Heitz, A. and Jung, Peter 2001. Plio-Pleistocene Faunal Turnover in the Caribbean: Molluscan Food Webs Implicate Plummeting Nutrient Supply. ,


Fortunato, Helena M. 2000. Larval Development of the Strombina-Group and Related Gastropods: Calibration of the Morphological Variation of the Larval Shell for Inference in Fossil Species. , : 13..

Fortunato, Helena M. 2000. Neogene Marine Biota of Tropical America (NMITA): World-Wide Web Taxonomy Database. , : 149..


Fortunato, Helena M. 1999. Biogeography and the Tempo of Speciation in Strombinid Gastropods. , : n...

Jackson, Jeremy B. C., Fortunato, Helena M., Todd, Jonathan A., Heitz, Antoine, Alvarez F., Marcos A., Johnson, Kenneth G. and Jung, Peter 1999. Molluscan Diversity Increased With Declining Productivity in Tropical America. , : A399..

Jackson, Jeremy B. C., Todd, Jonathan A., Fortunato, Helena M. and Jung, Peter 1999. Diversity and Assemblages of Neogene Caribbean Mollusca of Lower Central America. , : 193 -230..


Fortunato, Helena M. 1998. Reconciling Observed Patterns of Temporal Occurrence With Cladistic Hypotheses of Phylogenetic Relationship. American Malacological Bulletin, (2): 191-200..

Fortunato, Helena M. and Jackson, Jeremy B. C. 1998. Contrasting Modes of Larval Development Across the Isthmus of Panamá. , : 104..

Fortunato, Helena M., Penchaszadeh, Pablo E. and Miloslavich, Patricia 1998. Observation on the Reproduction of Bifurcium Bicanaliferum (Sowerby, 1832) (Gastropoda: Columbellidae: Strombina-Group) From the Pacific Coast of Panamá. Veliger, (2): 208-211..


Budd, Ann F., Foster, Jr, Charles T., Golden, Julia, Dawson, John P., Chidsey, Jennifer L. and Fortunato, Helena M. 1997. Surfing fossil biodiversity data on the World-Wide Web; the Neogene marine biota of tropical America (NMITA) database. , : 103-104..

Cavalier, Jaime, Jaramillo, Marie, Solis, Daniel and De Leon, Doris 1997. Water balance and nutrient inputs in bulk precipitation in tropical montane cloud forest in Panama. Journal of Hydrology Amsterdam, 193(1-4): 83-96..

Fortunato, Helena M. 1997. Calibrating phylogenies with the fossil record. Annual Report Western Society of Malacologists, 30: 21-22..

Fortunato, Helena M. 1997. Desarrollo larval en el grupo Strombina: calibración de la variación morfológica del protoconcho y su utilización para inferir el tipo de desarrollo en especies fósiles. , : 110..


Ashe, James S. and Lingafelter, Steven W. 1996. Revision of Gansia sharp of Mexico and Central America (Coleoptera: Staphylinidae; Aleocharinae). Journal of the New York Entomological Society, 103(3): 251-280..

Cavelier, Jaime, Solis, Daniel and Jaramillo, Maria Alejandra 1996. Fog interception in montane forests across the Central Cordillera of Panama. Journal of Tropical Ecology, 12(3): 357-369..

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